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Issue 21, 2026

Online ISSN: 2466-4367

, Issue 21, (2026)

Published: 22.01.2026.

Open Access

This Special Issue of Reforesta, International Practices for Regenerating and Restoring Forest Trees by Seeding, brings together regional practice and research from North America, South America, Europe, Asia, and Oceania on using broadcast and direct seeding to create forest cover. It presents a significant range of seeding activities inclusive of species, forests, sites, and climates. Seeding has regained interest because of its scalability and cost-effectiveness, particularly for remote or inaccessible areas, sites with low productivity where planting seedling costs are prohibitive, or when aiming for a more natural forest structure. Technological innovations, particularly in seed treatments, automation, and precision seeding improve success of large-scale seed dispersal.

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Contents

26.12.2016.

Research paper

Response of rare and endangered species Picea omorika to climate change - The need for speed

Serbian spruce (Picea omorika (Pančić) Purk.) is a rare and endangered tertiary relict and endemic species, with restricted and fragmented natural range in Serbia and Bosnia and Herzegovina, mainly around the mid-course of the Drina river. Since the middle of the 19th century, its natural range declines constantly, followed by a decline in the number of mature individuals. The decline of this forest species is slow and mainly attributed to poor regeneration and low competing ability. Given the foreseen worsening of the climate in forthcoming decades, this decline can only accelerate. In recent years, dieback related to drought has been observed as response to extreme weather events suggesting that Serbian spruce will face difficulties in adapting to climate change within its natural range. However, successful use of Serbian spruce in Central and Northern Europe indicates potentially large adaptive potential of this species which, along with the high genetic variability, outweigh the limited morphological variation, self-fertilization, and limitations related to the restricted natural range in the first place, and, indicates possible directions of migration in the second place. In this paper, current conservation actions are discussed, and strategies for the species survival in a changing environment are suggested. Since migration and adaptation are the least likely responses of this species to climate change, measures such as assisted migration may be the only strategy which will enable persistence of Serbian spruce. Current conservation programs, limited to in-situ actions, need to be supplemented with ex-situ actions and strategies. In the worst case scenario, i.e. for species such as Serbian spruce which are unable to migrate and/or adapt to changing climate, the most suitable sites should be identified and colonized in order to prevent extinction in the near future.

Vladan Ivetić, Jelena Aleksić

26.12.2016.

Research paper

Seed Source Effect on Quality and Morphology of Turkish Red Pine (Pinus brutia Ten.) Seedlings

This study was carried out to compare quality and morphology in 1+0 year containerized seedlings of Turkish red pine (Pinus brutia Ten.) originating from a seed stand and a seed orchard based on height and root-collar diameter. Averages of seedling height and root-collar diameter were 13.8 cm and 2.63 mm in orchard seedlings, while they were 14.8 cm and 4.56 mm in stand seedlings, respectively. There were significant differences (p≤0.05) between seed sources for the characters according to result of ANOVA. Stand seedlings were better quality than that of orchard seedlings according to quality classification of Turkish Standard Institute. Positive and significant (p≤0.05) relations were found between the characters in both seed sources based on results of correlation analysis. Results of the study were discussed for nursery practices, plantation forestry, and genetic-breeding of the species. 

Mahmut Cercioglu, Nebi Bilir

25.05.2016.

Review paper

Reforestation challenges in Southeast Europe facing climate change

Projections of the regional climate model for Southeast Europe generally predict an increasing of temperature and a decrease in precipitation, with some local variations. Higher frequency of extreme weather events and increased flooding can also be expected. This climate change will, among other things, result in changes in habitats and species distribution, and a decrease in biodiversity. In most cases, forest ecosystems will be unable to adapt fast enough to keep pace with changes in climate. Extreme weather events and low precipitation during the growing season will cause high mortality of seedlings after planting. New forests will face the whole range of these changes because of the long lifetime of trees. Reforestation programs must take projections of climate change into consideration. In the long term, new guidelines for site-species matching, provenance selection, and genetic diversity need to be adopted. In the short term, site preparation, planting techniques, and post planting protection need to be improved. In addition, seedling quality (morphological, physiological, and genetic) and planting time need to be specific for each site. New site preparation, planting, and post-planting protection methods are useful tools for short term success measured in seedling survival and initial growth. Seedling quality is essential for short and long term success. Different strategies, such as assisted migration and increased genetic diversity of planting material, can provide better chances for long term success measured in growth, fitness, and capability to produce the next, better adapted generation.

Vladan Ivetić, Jovana Devetaković

25.05.2016.

Review paper

Restoration Silviculture: An Ecophysiological Perspective - Lessons learned across 40 years

Involvement in forest restoration programs across North America for the past 40 years, dealing with nursery cultural practices, operational seedling quality programs and defining seedling performance on restoration sites has given me a unique perspective, which I have used to examine programs from both a research and operational perspective. Certain biological patterns and themes continually appeared across these programs and this paper discusses five of the most common themes.

Learning To Think Like a Tree – It is important for practitioners to develop an understanding of the ecophysiological performance of tree species in a nursery or forest restoration program in order to understand how seedlings grow. This understanding leads to sound biologically based cultural decisions to improve seedling performance.  

Stress and the Cyclical Nature of Stress Resistance – Seedlings are exposed to stress when environmental conditions limit their performance. Plants develop physiological resistance attributes to mitigate stress and these attributes change throughout the seasonal cycle. Practitioners have developed hardening cultural practices that enhance seedling stress resistance, thereby improving seedling quality and site restoration success.

Seedling Quality: Product versus Process – Seedling quality is an important component of successful restoration. Typically seedling quality is examined from a product perspective, thus defining functional integrity, operational grading or sometimes performance potential. An alternative approach monitors the process, with product quality the final output.

Planting Stress and Seedling Establishment – Planting stress is prevalent in forest restoration. The act of planting can result in a seedling that does not have proper connections for water movement through the soil-plant-atmosphere continuum (SPAC). Seedling water stress, reduced growth performance and potentially death can occur if this SPAC connection is not restored.  

Seedling Death: Sometimes Simple and Sometimes Complicated – Seedling death can occur in restoration programs as a result of environmental extremes or incorrect management practices. Some problems can be easy to diagnose and correct practices can be implemented to rectify the problem. Other times, issues are complicated and it can be a challenge to define the potential factors causing seedling death.  

Steven C Grossnickle

26.12.2016.

Research paper

Initial height and diameter are equally related to survival and growth of hardwood seedlings in first year after field planting

This study compares the relation of initial height and root collar diameter of bareroot hardwoods seedlings to survival and growth in first year after planting. A total of six species used in Serbian reforestation programs were tested (four native: Fagus sylvatica, Ulmus laevis, Fraxinus excelsior, and Acer pseudoplatanus and two exotic: Robinia pseudoacacia, and Quercus rubra), at 6 sites with a wide range of environmental conditions. Initial seedling height and diameter were equally related to field performance and better in forecasting growth than survival. The relation between seedlings initial morphological attributes and survival was species specific, while for all tested species growth was positively correlated to seedling size at planting. Although large seedlings kept their advantage in size, smaller seedlings grew at a higher rate. Both initial H and D should be considered as equally important in operational programs for hardwoods seedling quality testing.

Vladan Ivetić, Jovana Devetaković, Zoran Maksimović

25.05.2016.

Review paper

Review of the most important pathogens in Serbian forest nurseries

Successful reforestation depends on the quality and health of seedlings. Targeted production in forest nurseries should produce plants that will be able to survive unfavorable environmental and weather conditions in Serbia. Pathogens can reduce the vitality of seedlings and decrease survival after outplanting. The most common pathogens identified on seeds of both conifer and broadleaved seedlings are presented. The most frequently used fungicides for the control of pathogens in Serbian forest nurseries, as well as the best time of their application, are reviewed.

Nenad Keča

26.12.2016.

Review paper

Management and restoration of Italian afforestations under global change

As a consequence of past deforestation, degraded areas in Italy have been subjected to afforestation since the beginning of the XXI century. These afforestations have been done mainly with conifers (Pinus nigra, Pinus pinaster, Pinus halepensis, Pinus pinea), and with the aim to 1) protect the soil and to 2) prevent floods. Exotic forest tree species have been used for the afforestation of selected sites of the Apennine. In the case of Douglas fir, it is possible to state that after ninety years the results are very good in terms of both ecological adaptability and high growth-rate.

Nowadays, the lack of silvicultural treatments, the ageing processes, insect and fungi outbreaks have led pine afforestations to a condition of an unsteady biological equilibrium in many sites. Moreover, this condition seems to be due to additional factors related to climate change such as wind storms and dryness. The restoration of these pine afforestations is therefore one of the main aims of the present Italian forest management policy which aims to increase their level of resilience. In particular, the priority of the researchers has been given to the most fragile stands where three possible objectives of restoration measures should be considered:

1. To rehabilitate conifer stands by introducing native broadleaves when these stands are in a very degraded condition, and when the local wood energy chain needs to be started;

2. To conserve the conifer stands in the cases where the cultural, aesthetic and recreational functions, are the prevailing obtainable ecosystem services;

3. To foster the mixed stands, with the conifers of the old cycle and native broadleaves, which could increase resilience to the extreme events.

In regards of Douglas-fir stands, new silvicultural models have been developed which aim to a) conserve these stands and to b) combine a higher growth rate with a much improved mechanical tree stability.

Orazio La Marca, Roberto Mercurio, Leonardo Nocentini

25.05.2016.

Review paper

Afforestation on bare lands – example of Ibar Gorge, Serbia

The afforestation of bare lands, sandstone, and skeletal terrains are one of the biggest challenges forestry scientists face. These terrains are characterized by specific ecological conditions that are generally unfavorable for the growth of woody species. These are usually shallow soils, unstable, and poor in nutrients and moisture. The characteristics of these habitats make said terrains unfavorable for the regeneration of forest vegetation. It is therefore crucial for the success of afforestation to gain detailed knowledge and understanding of environmental conditions. Only after the detailed research and study of field conditions can the selection of species for afforestation, including selection of species characteristics and technology of planting, begin. Mistakes made during previous establishments of green areas are one of the main reasons some species of vascular flora have disappeared. This alone expresses the undeniable importance of knowledge on habitat specifics, work schedule, and selection of species for afforestation. With the aim to implement the future afforestation within the planned scope, it is necessary to organize an effective nursery production of seedlings with characteristics that will suit the environmental conditions of bare lands, sandstones, and skeletal terrains whose afforestation is planned.

Vladan Popović, Tatjana Ćirković-Mitrović

25.05.2016.

Letter to the Editor

Opening Letter

"The best time to plant a tree is twenty years ago. The second best time is now." Chinese proverb There are five million hectares of new forests planted each year, according to Global Forest Resources Assessment (FAO 2015). How successful are we in planting these new forests? What are challenges that practitioners meet today in planting forests? What are challenges that forests established today will face in decades to come? There is a myriad of research results and a body of knowledge large enough to understand principles behind the field performance of planted forests. The environment is dynamic and inputs and outputs continually change, so there is a constant need for new research ranging from the global to microsite scale, and from the ecosystem and species to the population and genotype scale. The journal Reforesta offers a new site for publishing research results, presenting experiences, and bringing forward novel ideas and discussions on reforestation issues. Reforesta editors will strive to combine academic excellence with professional relevance and wish to appeal to both the professional and academic communities. There are a large number of scientific journals, many of which deal with forest ecosystems. So a reasonable question could be: Do we need yet another professional journal? We say yes, because of the following reasons. Reforesta will be a "niche" journal, focusing on research stemming from sourcing seed, to nursery cultural operations, to field planting (e.g., afforestation, reforestation and forest restoration). In addition, Reforesta will cover topics of forest genetics, tree breeding, and stand silviculture. We will encourage the submission of papers that provide interest and value to the international readership dealing with planted forests, such as: 1) novel ideas or approaches to reforestation challenges; 2) connections between plant ecophysiology and seedling field performance; 3) testing of new techniques and products in production of forest reproductive material, and at the planting site; and 4) reviews and discussions on timely and important topics. Establishment of the new journal can be compared to establishing a new forest. Initiation of the Reforesta journal will be supported by the editorial team (i.e. planning and site-species or site-provenance matching). The team will then select appropriate papers from the contributions of willing authors (i.e. selection of planting material, nursery operations, and culling). Then the stage is set: the web-site is operational; the journal is registered and applied for indexing (i.e. site preparation, monitoring and feedback). The inaugural issue is published (i.e. seedlings are planted) and the research community is notified (i.e. post-planting silviculture operations). The parallel between Reforesta and establishing a new forest will continue. The future success of Reforesta will be dependent on the interaction with its contributors and readership. The future size of the Reforesta audience will be dependent on its ability to secure its niche, and to provide services to the professional forestry community, while developing a symbiotic relationship with other forestry journals (i.e. competition and/or facilitation). Performance of Reforesta will be measured by monitoring readership and response levels (i.e. reforestation success) and its lifespan will be defined by its resilience and adaptation potential. Reforesta will follow the open access policy of non-profit journals, with no processing charges applied to any accepted articles and no fees for accessing articles published by Reforesta. This principle ensures that forest research is free and available to forestry professionals in support of a greater global exchange of knowledge. Like planted forests, Reforesta hopes to grow in an open environment with appropriate tending from the editorial team, thereby growing with needs of the professional forestry community. The Reforesta journal has one additional advantage. It is established solely on the enthusiasm of the editorial team. As long as there are enthusiasts recognizing the need for Reforesta, the journal will continue to flourish. The editorial team is here to set the scientific standards and ethical rules for the journal. However, the ultimate direction of Reforesta depends on authors and readership. We are delighted to launch this new journal, a product of a joint venture between enthusiasts ranging from undergraduate students to senior researchers and scientists. Reforesta has one underwriting sponsor - The Scientific and Professional Society Reforesta. We thank them for their support. Under the leadership of Vladan Ivetić, Editor-In-Chief, who initiated the idea for Reforesta, and Steven Grossnickle, Consultant-Editor who plays a supporting role; we have a worthy team. The inaugural issue of the Reforesta journal is in form of Thematic Proceedings resulting from the International Conference on Reforestation Challenges, which took place on 3-6 June 2015 in Belgrade, Serbia. We wish to thank participants who attended the Conference and especially thank the invited authors who contributed to this inaugural issue. We are also grateful to the technical team, made up of excellent young students and researchers. We will continue to develop and fine-tune content to enhance the quality, scope and diversity of Reforesta as the journal grows into an established member of the professional forestry community. 

Vladan Ivetić, Steven C Grossnickle

26.12.2016.

Research paper

Height growth characteristics of one-year-old northern red oak seedlings (Quercus rubra L.) in full light conditions

In this nursery experiment, 100 one-year-old northern red oak seedlings were measured at the end of the 2016 growing season. Measured growth elements were: total height, root collar diameter and total leaf area. All seedlings were classified according to the presence of polycyclism into four types: one-flush growth, two-flush growth, three-flush growth, and four-flush growth type. The number of new flushes (shoots) is a good indicator of total height, root collar diameter and total leaf area. 

Nikola Miroljub Šušić, Martin Bobinac, Ivona Kerkez, Andrijana Bauer Živković, Nikola Vojinović