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Issue 21, 2026

Online ISSN: 2466-4367

, Issue 21, (2026)

Published: 22.01.2026.

Open Access

This Special Issue of Reforesta, International Practices for Regenerating and Restoring Forest Trees by Seeding, brings together regional practice and research from North America, South America, Europe, Asia, and Oceania on using broadcast and direct seeding to create forest cover. It presents a significant range of seeding activities inclusive of species, forests, sites, and climates. Seeding has regained interest because of its scalability and cost-effectiveness, particularly for remote or inaccessible areas, sites with low productivity where planting seedling costs are prohibitive, or when aiming for a more natural forest structure. Technological innovations, particularly in seed treatments, automation, and precision seeding improve success of large-scale seed dispersal.

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30.06.2017.

Research paper

Using double-sampling techniques to reduce the number of measurement trees during forest inventories

Variable-radius sampling techniques are commonly used during forest inventories. For each sample tree at a particular sampling point, diameter and height(s) are measured and then weight is estimated using established equations. Heights can require a fair amount of time to measure in the field. Separating the weight per acre estimate into two components; average basal area per acre and WBAR (individual tree weight-basal area ratio) across all points, can often lead to more efficient sampling schemes. Variable-radius sampling allows for a quick estimate of basal area per acre at a point since no individual tree measurements are needed.  If there is a strong relationship between weight and basal area, then by knowing basal area you essentially know weight. Separation into two components is advantageous because in most cases there is more variability among basal area estimates per point then there is in WBAR. Hence, you can spend more resources establishing many points that only estimate basal area – often called “Count” points. “Full” points are those where individual tree measurements are also conducted. There is little published information quantifying the impacts on basal area, weight, etc., estimates among different “Full/Count” sample size ratios at the same site. Inventories were examined to determine this method’s applicability to loblolly pine plantations in southern Arkansas and northern Louisiana. Results show there is more variability among basal area estimates than WBAR and that the amount of trees being “intensively” measured is excessive.  Based on these four plantations, a “Full” point could be installed ranging from every other point to every fifth point depending on site conditions and the desired variable.

Curtis L. VanderSchaaf, Gordon Holley, Joshua Adams

30.06.2017.

Research paper

Should forest regeneration studies have more replications?

When it comes to testing for differences in seedling survival, researchers sometimes make a Type II statistical error (i.e. failure to reject a false null hypothesis) due to the inherent variability associated with survival in tree planting studies. For example, in one trial (with five replications) first-year survival of seedlings planted in October (42%) was not significantly different (alpha = 0.05) from those planted in December (69%). Did planting in a dry October truly have no effect on survival? Authors who make a Type II error might not be aware that as seedling survival decreases (down to an overall average of 50% survival), statistical power declines. As a result, the ability to declare an 8% difference as “significant” is very difficult when survival averages 90% or less. We estimate that about half of regeneration trials (average survival of pines <90%) cannot declare a 12% difference as statistically significant (alpha = 0.05). When researchers realize their tree planting trials have low statistical power, they should consider using more replications. Other ways to increase power include: (1) use a one-tailed test (2) use a potentially more powerful contrast test (instead of an overall treatment F-test) and (3) conduct survival trials under a roof. 

David B. South, Curtis L. VanderSchaaf

30.06.2017.

Review paper

Afforestation in Bulgaria

The historical development of afforestations in Bulgaria was investigated. Some of the more important decisions and events which determine the correct state policy in the field of afforestations are pointed out. The most significant achievements are in: erosion control, incl. water tank protection from silting up; increasing the tree volume and forest growth; protection of agriculture via forest protection belts; landscape improving. The social importance for more employment and improvement of means of livelihood is mentioned. Proved reasons for decreasing the afforested areas since 1980s are given. The main mistakes and problems are mentioned as well as the changes in the afforestation policy and future challenges.

Milko Milev, Krasimira Petkova, Nasko Iliev

30.06.2017.

Research paper

Application and use of sulphuric acid pretreatment to improve seed germination of three acacia species

The experiments were led to study the effect of pretreatments and their duration on germination behavior of three Acacia species A. cyanophylla Lindl., A. farnesiana L. and A. decurrens Willd. by analyzing three parameters (GP: germination percentage; MGT: germination mean time (days) and GRI: germination rate index) for various times of incubation (5, 10 and 15 days) in Petri dishes. Pre-sowing treatment included immersion in concentrated sulphuric acid for 60, 90 and 120 minutes. The sowing in distilled water (Control) had no positive effect on the germination induction. Generally, the seed pretreatments were very useful to improve germination. Time of immersion significantly (P < 0.0001) affected GP, MGT and GRI in all species. Increasing the duration of sulphuric acid immersion (from 60 to 120 min) improved the germination percentages for A. cyanophylla and A. farnesiana seeds to (98% and 99%), respectively. However, increasing this duration had a negative effect on A. decurrens seed germination, reducing the final germination percentage from 97% at 60 minutes of immersion to 43% at 120 minutes.

Abdenour Kheloufi, Lahouaria Mounia Mansouri, Faiza Zineb Boukhatem

30.12.2017.

Research paper

Sulfur and lime affect soil pH and nutrients in a sandy Pinus taeda nursery

Two pH experiments were conducted at a sandy, bareroot loblolly pine (Pinus taeda L.) nursery in Texas. A sulfur trial (0, 813, 1626, 2439 kg ha-1 of elemental sulfur) was installed to determine if lowering soil pH would result in nutrient toxicity symptoms and affect seedling morphology. Although soil acidity in the sulfur study ranged from pH 3.9 to pH 5.0, none of the treatments resulted in micronutrient toxicity and none affected height growth, root-collar diameter, root mass, shoot mass or the root-mass ratio (root dry mass/total dry mass). Acidifying soil with sulfur increased leaching of calcium, potassium, magnesium, manganese and zinc but there was no effect on seedling morphology. The objective of the liming trial (0, 813, 1626, 3252 kg ha-1 of dolomitic lime) was to determine if increasing alkalinity would result in an iron deficiency and reduce seedling growth. As expected, applying lime increased the calcium and magnesium levels but had no effect on soil levels of iron, phosphorus, potassium, sulfur, zinc and sodium. However, the root-mass ratio was reduced by applications of dolomitic lime (pH ranged from 5.3 to 6.0). Differences in soil properties (i.e. plot location) had a greater effect on seedling morphology than lime applications. Foliage levels of manganese and boron were reduced by the highest rate of lime and sulfur, respectively.

David B South, Ryan L Nadel, Scott A Enebak, Gene Bickerstaff

30.06.2017.

Review paper

Reforestation in Poland: History, Current Practice and Future Perspectives

In the past, the entire region of Poland was overgrown by forests. Due to economic changes, the forest cover was reduced to 40% in the 18th century and 21% after the Second World War. After the war, Polish foresters undertook considerable efforts to increase the forest cover to 30.8% by 2015. Polish forests are characterized by the dominance of oligo- and mesotrophic coniferous species (68.7%). This include the pioneer species, Scots pine. It covers approximately 60% of the area. The species composition of Polish forests determined the dominance of artificial regenerations. However, the currently prevailing direction of forest culture is natural regeneration. This tendency is related to “greening” of the forest management, the priority of PERSONAL durability over productivity and culture of multifunctional forests. A natural or seminatural direction of forest culture is being promoted. Renewal of the species such as fir, beech, oak, or spruce from the last stages of succession have always taken place in a natural manner, whereas the statistics are generated by the dominant species preferring open areas during renewal. Currently, the scale of natural regenerations of the pine is increasing. It is increasingly common to value the favorable economic aspect of natural renewal of the species, and the experience of practitioners supported by scientific research increase the likelihood for success. In Poland, the majority of methods of regeneration proceedings (forest cutting) and the law are directed at obtaining and promoting natural renewal. Independent of the concept of natural renewal promotion, the location of Poland in the intermediate climate zone, between the influence of oceanic and continental climates, resulted in the formation of valuable tree stands with high flexibility and tolerance to growth conditions. They are divided into seed stands, excluded stands, and timberlands. Thus, Poland is in possession of a great base for seed collection. At the beginning of 1990s, a rapid need for container seedlings occurred due to numerous disasters (wind-broken trees, gradations). Currently, in Poland, 17 field nurseries are in operation producing 1–10 million seedlings. In 1992, Poland received a loan from the World Bank to conduct afforestations and the “National Program for Increasing the Forest Cover” was started. The main objective of the plan is to increase the forest cover to 30% in 2020 and 33% in 2050. Within the program, it is planned to include vegetation of the natural succession in the area of approximately 80,000 ha. 

Jacek Banach, Kinga Skrzyszewska, Jerzy Skrzyszewski

30.06.2017.

Research paper

Nutritional diagnoses of oriental beech trees in damaged Caspian forest sites, using the diagnosis and recommendation integrated system (DRIS)

Oriental beech established in the Hyrcanian forests, is a valuable tree whose habitats are constantly exposed to destructive factors which change the nutritional status of soil and leaves. Analysis of foliar elements is a commonly used method for studying tree nutrient status that indicates site's quality. Foliar analysis of beech (Fagus orientalis Lipsky) was carried out in Kojour (Mazandaran) in order to assess the nutritional balance of trees in a damaged forest site (a direct result of livestock grazing and anthropogenic perturbations). Sunny leaves of dominant trees were taken in August and foliar concentration of macroelements, N, P, Ca, K and Mg were measured. The diagnosis and recommendation integrated system (DRIS) analysis was applied for evaluating the nutritional state. The results showed deficiency with K and P in disturbed stands. Nutrient Balance Index (NBI) indicated imbalance in nutrient status. These results suggest the usefulness of DRIS for foliar tissue analysis as an indicator of nutritional status and elemental stress in natural forests.

Afrooz Alimohamadi, Hamid Hamid Jalilvand, Moslem Akbarinia, Abas Banje Shafiei, Khadijeh Afzali

30.06.2017.

Review paper

Concerns and evidence on genetic diversity in planted forests

Planted forests are an important source of various services (economic, environmental, and social) with increasing portion in the total world’s forest area. Genetic diversity is fundamental for success and sustainability of planted forests. Facing the concern of the reduction of genetic diversity in planted forests, this study offers a review of evidence on comparisons between the levels of genetic diversity in forests established by different regeneration methods. A total of 34 papers comparing genetic diversity in natural forests versus various regeneration methods of 24 tree species examined by the range of markers are reviewed. In most cases, there are no significant differences in genetic diversity between natural and planted forests, followed by an almost equal number of cases with decreased and increased level of genetic diversity. The loss of rare alleles, but also new alleles are reported in planted forests. Although the origin of planting material in the most cases are unknown, the size of parental population is determinant for the level of genetic diversity in the new forest, with the provenancing and seed collection strategy as the most important management practices in planting projects.

Vladan Ivetić, Jovana Devetaković

30.12.2017.

Review paper

Direct Seeding in Reforestation – A Field Performance Review

Direct seeding has been considered a forest restoration option for centuries. Over the past half century, the use of this practice has declined in developed countries as forest regeneration programs have advanced with the production of quality seedlings that can successfully establish restoration sites. Direct seeding is being reconsidered as a restoration option as the potential size of the worldwide forest restoration program has grown because of massive deforestation in third-world nations and due to global climate change. This review examines direct seeding from a number of perspectives. First, merits of using this practice in restoration programs are defined. Major merits of this option are that it can be done quickly, over hard to reach and large disturbed areas, and at a relatively low cost. Second, current research findings from restoration programs are discussed. The major finding is that seedling establishment rates are low (i.e. typically around 20% of seeds planted) due to site conditions, seed predation and vegetation competition, and field performance (i.e. survival and growth) is lower than planted seedlings. Third, operational practices for the application in restoration programs are reviewed. To successfully conduct direct seeding programs practitioners need to consider seedbed receptivity, seed distribution and seeding rate. Fourth, potential new practices are presented. Some of these new practices attempt to create a more effective means to disperse seed across the site, minimize seed predation or create a more favorable microsite environment. This review provides a synthesis of what is known about direct seeding, thereby allowing practitioners to make a rational decision of whether to apply this practice towards their forest restoration program.

Steven C Grossnickle, Vladan Ivetić

30.06.2017.

Review paper

Reforestation in Nigeria: History, current practice and future perspectives

Nigeria, a country in West Africa, sits on the Atlantic coast with a land area of approximately 90 million hectares and a population of more than 140 million people. The southern part of the country falls within the tropical rainforest which has now been largely depleted and is in dire need of reforestation. About 10 percent of the land area was constituted into forest reserves for purposes of conservation but this has suffered perturbations over the years to the extent that what remains of the constituted forest reserves currently is less than 4 percent of the country land area. As at today about 382,000 ha have been reforested with indigenous and exotic species representing about 4 percent of the remaining forest estate. Regrettably, funding of the Forestry sector in Nigeria has been critically low, rendering reforestation programme near impossible, especially in the last two decades. To revive the forestry sector government at all levels must re-strategize and involve the local communities as co-managers of the forest estates in order to create mutual dependence and interaction in resource conservation. 

Isidore Nelson Akpan-Ebe