Current issue
Issue 21, 2026
Online ISSN: 2466-4367
, Issue 21, (2026)
Published: 22.01.2026.
Open Access
This Special Issue of Reforesta, International Practices for Regenerating and Restoring Forest Trees by Seeding, brings together regional practice and research from North America, South America, Europe, Asia, and Oceania on using broadcast and direct seeding to create forest cover. It presents a significant range of seeding activities inclusive of species, forests, sites, and climates. Seeding has regained interest because of its scalability and cost-effectiveness, particularly for remote or inaccessible areas, sites with low productivity where planting seedling costs are prohibitive, or when aiming for a more natural forest structure. Technological innovations, particularly in seed treatments, automation, and precision seeding improve success of large-scale seed dispersal.
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Contents
29.06.2024.
Research paper
Enhancement of seed germination rate and growth of Anagyris foetida L.
Bean trefoil (Anagyris foetida L.), the only species within the Anagyris genus (L.) of the Fabaceae family in Algeria, demonstrates considerable promise for reforestation actions in arid and semi-arid areas throughout the Mediterranean basin. Nevertheless, a prominent obstacle impeding the successful establishment of the species is the hardness of its seeds, resulting in reduced germination rates and inconsistent seedling emergence. The objective of this research is to improve seed germination and seedling development in A. foetida. A total of 750 pods were randomly collected from a population including 13 individuals. From this sample, 100 intact pods and 100 seeds were chosen for measurement and weight assessment. Three different scarification techniques were utilized as pretreatments: chemical scarification, involving varying durations of immersion in 98% concentrated sulphuric acid; thermal scarification, involving 24 hours of exposure to a water bath set at 35°C; and mechanical scarification, achieved through the use of abrasive paper. Each treatment comprised four replicates of 50 seeds, while untreated seeds were utilized as controls. Germination and initial seedling development were assessed after 30 days of cultivation. Statistical analysis revealed significant differences (p<0.001) among pretreatments concerning germination and seedling emergence. The highest germination percentages were observed following 120 minutes and 150 minutes of sulphuric acid soaking, with 86% and 91.5% of germination, respectively. The chemical pretreatments demonstrated correlations with shoot and root growth, as well as collar diameter. In contrast, the control group exhibited notably low germination (5.5%), while mechanical scarification resulted in a 55% germination rate. Seedling survival rates ranged from 93.7% to 99.4%. The seeds of A. foetida predominantly exhibit physical dormancy attributed to their hard and impermeable seed coat. These findings could be valuable for the generative reproduction of the species, especially for producing plants for afforestation/reforestation programs.
Lahouaria Mounia Mansouri, Abdenour Kheloufi, Mohamed Boukhecha
29.06.2024.
Research paper
The effectiveness of agroforestry on agricultural productivity and erosion control in Rwanda
Agroforestry bridges the gap that often separates agriculture and forestry by building integrated systems that address both environmental and socio-economic objectives. Agroforestry can improve the resiliency of agricultural systems and mitigate the impacts of climate change. Existing research suggests that integrating trees on farms can prevent environmental degradation, improve agricultural productivity, increase carbon sequestration, generate cleaner water, and support healthy soil and healthy ecosystems while providing stable incomes and other benefits to human welfare. This research aims to provide for identifying and adopting the evidence demonstrating the effects of agroforestry practices and interventions on agricultural productivity, erosion control, ecosystem services, and human well-being. To study the effectiveness of agroforestry on agricultural productivity and erosion control in Rwanda, raw data was collected with the aid of a questionnaire and focus groups. It was found that most farmers (95.6%) adopted and were fully engaged in agroforestry while all the officials made up to the mark. Woody species like Grevillea robusta, Cassia siamea, Albizia spp, Acacia albida, Persea americana, and Mangifera indica were found to be tree species the most integrated with crops in all the provinces of Rwanda. It was evident that farmers practice agroforestry to enhance the overall sustainability of farming systems by improving agricultural productivity and preventing erosion.
Boniface Byiringiro, Ujjwal Akshay, Claude Kayijuka, Babita Bharti
29.12.2023.
Research paper
Responses at the stand and tree level to ice storm injuries in beech forests in eastern Serbia
The paper presents the study on the condition and structural stability of European beech (Fagus sylvatica L.) stands after the 2014 ice storm disturbance. The research was conducted in three pure beech stands on the Rtanj mountain in eastern Serbia. The stand condition after the ice storm was analyzed using changes in taxation elements. The assessment of tree damage was done according to the ICP Forests methodology, while the structural stability of the stands and individual trees was analyzed using the slenderness coefficient. The obtained results indicate very heterogeneous responses of beech stands to the negative impact of the ice storm. Heterogeneity is not only expressed between stands but also within individual stands. The stands after the ice storm are characterized by significantly reduced values of production indicators, but also by satisfactory stability, considering that the trees from the understory suffered the most. Statistically significant differences in the stability of trees of different dimensions expressed through the slenderness coefficient were determined. Consequently, the stands have maintained an appropriate level of stability thanks to the survival and resistance of the dominant trees that are the bearers of functions in these stands (trees with dbh > 40 cm). This indicates the great potential of silvicultural treatments that can increase the stability and resistance of stands in areas with a high risk of such phenomena. In addition, repressive action in the form of salvage logging has a very important role with the aim of ensuring the continuity of stands after damage has occurred.
Branko Kanjevac, Miloš Račić, Jovan Dobrosavljević, Nikola Martać
29.12.2023.
Research paper
Effects of Eucalyptus species on soil physicochemical properties in Ruhande Arboretum, Rwanda
This study combines research on soil physical and chemical properties as affected by four Eucalyptus species in Ruhande arboretum. The soil samples for research properties were taken from 0-20 cm depth using auger and one undisturbed core from each sampling unit was taken for the analysis of soil bulk density. Soil bulk density and moisture did not differ significantly between all treatments. Sand proportions differed significantly only between E. tereticornis and E. maidenii whereas silt and clay were non-significant. The soil under Eucalyptus was sandy, with sand proportion ranging from 66.4-71%. Bulk density increased with increasing sand whereas moisture content showed a reverse trend. The soil samples for studying chemical properties were taken as described in Nsabimana et al. 2008. All chemical parameters except base saturation differed significantly between treatments. Soil pH was strongly acidic but rich in total nitrogen and organic carbon which was attributed to higher litter production, its relatively faster rate of decomposition, and greater amount of residues produced by the eucalypts. Carbon/nitrogen ratio, CEC, and BS were high under all treatments while the available phosphorus was lower which was attributed to the low pH. Future studies should test if the species similarly affect the soil or not and confirm if the species increase soil nutrients. Benchmarked study sites should be used to enable differences in the species effects on the site if any.
Olivier Niyompuhwe, Charbel Maklouf Jabiro, Canisius Patrick Mugunga
29.12.2023.
Review paper
Zinc fertilization in bareroot pine seedbeds
Zinc deficiencies are rare in pine seedlings with less than five documented cases in bareroot nurseries. One temporary deficiency occurred after soil was land-leveled (i.e., topsoil removed) and another occurred on a peat soil after more than 2,200 kg of agricultural lime was applied before sowing. Farmers also observe zinc deficiencies on (1) over-limed areas and (2) where Zn-demanding crops are grown on areas where topsoil was removed during land leveling. Since ZnSO4 is a naturally occurring pesticide, sometimes height growth increases are due to pest control. In pathogen-rich soils, pine growth may be improved more by the fungicidal effect than by a growth benefit from added sulphur and zinc. As a result, a pseudo-deficient response is possible when growth of non-deficient seedlings increases after treatment with large amounts of ZnSO4 or ZnCl2. In some trials, claims of a Zn deficiency have been made without supporting evidence from foliar tests or from tests using pathogen-free soil. Although fertilization with Zn increased seedling growth at pine nurseries in New Zealand, India, Russia, and Wisconsin, only at the Sweetwater Nursery in New Zealand did foliar tests prove a Zn deficiency.
David B. South
29.12.2023.
Research paper
Monitoring of groundwater level fluctuations at flooded area of lowland forests of the Sava River (Serbia)
Lowland pedunculate oak forests in the alluvium of the river Sava are of a great economic and ecological importance. Their growth and development features, and their survival, as well, mostly depend on soil moisture and available water. The paper deals with the influence of groundwater level on pedunculate oak forests condition in flooded area of Donji Srem. During four-year long research, the average (reference) groundwater level was determined, and extremes, the wettest and the driest year, were determined based on climate factors such as temperature and rainfalls. Deviation of the average from the reference groundwater level in the vegetation period during some years indicates possibility of risk zones presence. These zones are defined according to geostatistical analysis – ArcGIS, Kriging model and they are very significant for forestry practice, because they can predict some drought events and warn forestry experts to react in time in order to prevent big damages. Bearing in mind that investigated area is flooded, and that groundwater level is close to the soil surface, these anaerobic site conditions are obviously more suitable for another hygrophilous species than for pedunculate oak.
Vesna Nikolić Jokanović, Andreja Lazović, Tihomir Šoškić, Kristina Živanović
04.07.2023.
Research paper
Variability of morpho-anatomical features of black pine needles at the area of Jastrebac and Goč
In this paper the variability of morphological (needle length and needle width) and anatomical (needle thickness, resin ducts width, epidermis thickness, hypodermis height and number of the hypodermis layers) properties of black pine (Pinus nigra J.F. Arnold) needles were studied at the intra- and inter-population levels. Two mountains in Serbia, Jastrebac and Goč, were selected as experimental plots. Three Pinus nigra trees were selected from both localities (6 in total). Obtained results showed that trees on Goč had bigger dimensions of the following elements: needle length, slightly wider needle width, epidermis thickness, hypodermis height, number of hypodermis layers and needle thickness. On the other hand, resin ducts were only slightly wider by the trees from Jastrebac. The results of the analysis of variance showed that variation between studied populations, as well as variation between trees within populations was statistically significant for all needle traits except resin ducts width and hypodermis height.
Kristina Živanović, Dušan Jokanović
04.07.2023.
Review paper
Use of sulphur in bareroot pine and hardwood nurseries
During the 20th century, managers at sandy nurseries utilized sulphur (S) to lower soil pH and mitigate the risk of iron deficiency. During that time, however, applying S as a fertilizer was a rare event. At many nurseries, S in rain and irrigation water was sufficient to avoid visual deficiency symptoms. The S status of soil and foliage was typically unknown, and many researchers did not test for S due to the additional cost. Consequently, S became the most neglected macronutrient. While a few nursery trials demonstrated that elemental S reduced damping-off and increased height growth, a majority showed no benefit after applying S at rates lower than 100 kg ha-1. Even so, by 1980, S-deficiencies occurred at bareroot nurseries in Alabama, Oklahoma, Virginia, Wisconsin, the United Kingdom, and likely in North Dakota and New York. The risk of a deficiency increases when N-only fertilizers are applied to seedbeds. Due to research, experience and the precautionary principle, several managers transitioned to using ammonium sulfate instead of, less expensive, N-only nitrogen fertilizers. After soil tests became affordable, managers began to ask questions about the need to apply S to seedbeds. Only a few hydroponic trials with small pine seedlings have been used to estimate “threshold” or “critical values” for foliar S. Since an initial 1,500 μg g-1 S value is “unreliable” for pine seedlings, some authors lowered the value to 1,100 μg g-1 and even as low as 500 μg g-1 S. Others ignore all estimates based on total S concentrations and, instead, monitor only foliar SO4 levels.
David B. South
29.12.2023.
Research paper
Assessment of carbon sequestration of Teak (Tectona grandis Linn. F.) plantation on the campus of University of Ilorin, Nigeria
The study assessed the carbon sequestration of Tectona grandis Linn. F. in five age series (11-15 years old), at the University of Ilorin, north-central Nigeria. Data were collected using a stratified sampling technique and twenty square plots of 25 m × 25 m were laid. A non-destructive method was used to determine the biomass of the trees. Tree enumerations were carried out for diameters at the base, top, middle, diameter at breast height (DBH), and height. Soil samples at two different depths (0-15 cm and 15-30 cm) were collected and analyzed to obtain soil organic carbon. The results were summarized using descriptive statistics, while the relationship between tree growth variables and carbon stock was assessed using correlation and regression analysis. The results showed that tree carbon stocks were 230.05 t ha-1, 362.35 t ha-1, 277.48 t ha-1, 216.40 t ha-1, and 126.20 t ha-1 for 11 years old (2012), 12 years old (2011), 13 years old (2010), 14 years old (2009) and 15 years old (2008) age series, respectively. The soil organic carbon stocks were 1.1025 t ha-1, 0.6253 t ha-1, 1.2019 t ha-1, 1.4070 t ha-1 and 0.7615 t ha-1 for 11 years old (2012), 12 years old (2011), 13 years old (2010), 14 years old (2009) and 15 years old (2008) age series, respectively. The study also revealed that the 14-year-old (2009) age series had the highest carbon stock, and the total carbon stock estimate was 151,850.84 t. The potential cash value of the carbon stock was also estimated at $91,894.40 t C ha-1 and the total PCV was $56,606,951.50. Correlation analysis showed a strong positive correlation between most of the growth variables and carbon stock. The regression equation (Y = -596.48 + 27.16 THT + 1238.34 DBH, Adj R2 = 82.7%) showed that DBH and height of trees are suitable for evaluating the carbon stock in the study area.
Tajudeen Okekunle Amusa, Mustapha AMINU, Farhan Moshood
29.12.2023.
Research paper
Implications of incorrectly determining site index on stand-level management activities and financial returns in older generation loblolly pine plantations
Predicting future yields normally requires an estimate of site quality. A commonly used measure is site index (SI). SI is often incorrectly quantified operationally due to the ambiguity associated with selecting “site” trees. Plus, error in the measurement of height itself occurs. This study quantifies the impacts on the number and timing of thinnings, and the final harvest ages, as well as financial returns when incorrectly determining SI. Three values of SI (base age 25 years) were examined using two older generation loblolly pine plantation growth and yield simulation models from the Western Gulf, USA; 16.76 m, 21.34 m, and 25.91 m. Firstly, a particular SI was assumed to be the “true” value, growth and yield estimates were obtained, and financial assessments were conducted. The same process was then conducted again, but assuming that the SI was incorrectly determined by varying positively and negatively the SI by up to 1.22 m from the assumed “true” value. For these older generation plantations, incorrectly determining SI did impact the age of the first thinning by as much as 5 years. In some cases, errors of +/- 1.22 m in SI estimation had little impact on the estimated timing of the first thinning. Errors in SI of up to +/- 1.22 m had little impact on the number of thinnings across economic rotation ages. For both unthinned and twice-thinned stands, final harvest (clearcut) ages differed by as much as 4 years for SI errors up to +/- 1.22 m. These errors led to differences in Land Expectation Value (LEV) up to $406.50 ha-1. Across the three SI (16.76, 21.34, and 25.91 m), differences in LEV ranged from $237.49 to $406.50 ha-1. These differences in LEV could be enough to incorrectly not conduct, or incorrectly conduct, a silvicultural operation such as an herbicide treatment or a fertilization treatment across a rotation, among other treatments.
Curtis VanderSchaaf