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Issue 21, 2026

Online ISSN: 2466-4367

, Issue 21, (2026)

Published: 22.01.2026.

Open Access

This Special Issue of Reforesta, International Practices for Regenerating and Restoring Forest Trees by Seeding, brings together regional practice and research from North America, South America, Europe, Asia, and Oceania on using broadcast and direct seeding to create forest cover. It presents a significant range of seeding activities inclusive of species, forests, sites, and climates. Seeding has regained interest because of its scalability and cost-effectiveness, particularly for remote or inaccessible areas, sites with low productivity where planting seedling costs are prohibitive, or when aiming for a more natural forest structure. Technological innovations, particularly in seed treatments, automation, and precision seeding improve success of large-scale seed dispersal.

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Contents

31.12.2019.

Review paper

Potassium fertilization in bareroot nurseries in the southern US: a review

This review covers most of the published literature on potassium (K) fertilization in bareroot seedbeds with the intent to concentrate on the southern United States. The timing and rates of K fertilization for bareroot seedlings are often based on logic and myths and, as a result, K recommendations vary considerably. Some recommend bareroot pine seedlings be fertilized with twice as much K as nitrogen (N) while others apply less than 100 kg ha-1. It was determined that several long-held claims about K fertilization are invalid. Nursery seedbeds do not need to contain four times as much available K as N and the belief that extra K fertilization will increase freeze tolerance or drought resistance of non-deficient seedlings is invalid. There are no data to support the claim that K fertilization increases root growth or assists in the formation of terminal buds. For sandy seedbeds, there is no need to apply K before sowing. Adding extra K during the fall does not increase seedling morphology or seedling performance when loblolly pine seedlings, at lifting, have more than 0.5% K in needles. A reduction of K fertilization can be achieved by reviewing foliar tests prior to K top-dressings.

David B. South

31.12.2019.

Review paper

Reforestation in Venezuela – current situation and future perspectives

The location of the Bolivarian Republic of Venezuela in the tropical region of northern South America, in the Northern Hemisphere, with a total area of 91.64 million ha, of which 49.6 million correspond to natural forests (54.12 % of the national territory). Venezuela shows a series of geographic advantages that allow a rapid growth and development of the trees in comparison with the temperate zones where the growth is extremely slow. However, the occurrence of deforestation processes has intensified in a sustained way in the recent history of the country.

In the period from 1920 to 2008, the loss of 26.43% of the forested area has been recorded. Likewise, in the period 1975-1988, deforestation was estimated at 216,000 ha year-1, equivalent to 2.9 million ha in 13 years (Catalan 1992). Deforestation has now increased to worrying levels and according to FAO estimates (2010), in Venezuela, a forested area of more than 288.000 ha year-1 is deforested.

In the middle of the 20th century several public and private institutions began an important effort to reforest areas of sandy soils of scarce depth on a bed of highly impermeable clay. With erosion processes generated as a result of strong winds that causes severe damage to the little existing vegetation. It was expected that with the passage of time this region could become a desert in the oriental plains of Venezuela. This is how the Maderas del Orinoco Companies (CVG - PROFORCA), the Venezuelan Corporation of Guayana (CVG), Smurfit Kappa Cartón of Venezuela, San Carlos Forestry Development, Masisa. Propulso, among others, started in different regions of the country the establishment of plantations for various purposes. For this they have used fast growing forest species such as: Pinus caribaea var hondurensis, Eucalyptus Urophylla, Acacia magnium, Caraipa llanorum, Tectonia grandis, Hebea brasiliens, Gmelina arborea, Erisma uncinatum, among others. At present, a reforested area of 650.000 ha has been reached. This has favored the protection of natural habitats by reducing the pressure for exploitation and use of the forests.

The legal framework established in the new National Constitution, approved in 1998, is based on decrees of 1825-1829 of the Liberator Simón Bolívar and classifies the territory in diverse forest areas establishing a modern environmental conservation policy. A historical summary and the current legislation with the classification that promises a rational management of the national territory is presented.

Francisco Visaez, Eduardo D. Greaves

31.12.2019.

Research paper

Application of a PhotoThermal model for container-grown conifer seedling production

This study applied a total energy approach to model seedling growth for container-grown loblolly pine (Pinus taeda L.). Seedlings were grown in three container stocktypes representing a range of cavity volume and density patterns. These seedlings were grown under both controlled greenhouse and outside compound environmental conditions under well-defined cultural conditions. Models for temperature and light ranges were created from work on the ecophysiological performance and morphological development of loblolly pine to these atmospheric conditions. A PhotoThermal data set was created by generating hourly averages of these two environmental variables during the growing season. Light and temperature data were integrated, each weighted equally, into PhotoThermal hours (PTH) to assess the crop growth response. Loblolly pine seedling growth in both the greenhouse and outside compound was directly related to PTH. Seedling growth was also related to the container type with the largest cavity volume and lowest cavity density having the greatest growth per PTH. Application of the PhotoThermal model is discussed for growing seedlings in an operational program having multiple production steps, delivery dates and nursery locations.

Steven Grossnickle

28.06.2019.

Research paper

Seed germination of Crataegus monogyna – a species with a stony endocarp

The present work demonstrates the effects of moist cold stratification on seed dormancy breaking in Hawthorn (Crataegus monogyna Jacq.). We also examined the fruit and seed morphology. Mature and ripe fruits were collected, the pulp removed and the seeds (stones) left in the sun to dry for three days. Four temperature regimes viz. 4°C, 6.5°C (natural conditions where the fruits were harvested), 10°C and 20°C were used for stimulating seed germination under total darkness. For each treatment, there were four replicates with 50 seeds incubated in a plastic container between two layers of moist sand at 15%. At the end of the experiment, non-germinating seeds were tested for viability using Tetrazolium chloride (TZ).  After 4 months, the final germination (FGP) was expressed as a percentage of the total number of seeds in each treatment. The fruits of Hawthorn were 14.9 ± 0.73 mm long and 15.1 ± 0.84 mm in diameter and weigh 2.05 ± 0.28 g. The seeds were 8.29 ± 0.43 mm long and 6.75 ± 0.39 mm in diameter and weigh (0.25 ± 0.04) g. The thousand-fruit weight was 2,000 g and of the thousand-seeds weight was 280 g. The statistical analysis indicated significant effect (p < 0.0001) of treatment on seed germination. Dormancy in this species was broken most effectively by cold stratification at 4°C and under natural conditions with 76% and 67.5% of FGP, respectively. Most of the nongerminated seeds of C. monogyna were viable (dormant) as judged by TZ.

Abdenour Kheloufi, Lahouaria Mounia Mansouri, Cécile Vanbellinghen

28.12.2018.

Research paper

The effect of seed size grading on seed use efficiency and height of one-year-old container-grown Scots pine (Pinus sylvestris L.) seedlings

The seed of Pinus sylvestris L. originating from natural forest in Voronezh region, Russian Federation, was graded in four classes based on seed diameter size, using the operational equipment of screens with round openings. The ungraded seedlot was used as control. Seedlings were produced in Voronezh containerized forest nursery, from March to September 2015, following the standard procedure for Pinus sylvestris. Seed from each size class, plus control, is sown in containers with 120 cm3 cells, filled with peat-substrate, grown in greenhouse for eight weeks, followed by hardening in growing area until the end of the growing season. The results shows a week positive effect of seed grading on height of one-year old container-grown seedlings, but significant improvement of seed use efficiency by reducing the number of empty cells. Based on results of this study, we can recommend grading of Pinus sylvestris seed on two size classes.

Arthur I. Novikov, Vladan Ivetić

10.07.2018.

Letter to the Editor

Reforestation challenges 2018

The conference on "Reforestation Challenges" took place on 20-22 June 2018 in Belgrade, Serbia, at Faculty of Forestry of the University of Belgrade. This was the fourth meeting on reforestation challenges, after three successful meetings in three previous years (2015 – Belgrade, Serbia; 2016 – Krakow, Poland; 2017 – Session at the IUFRO Congress in Freiburg, Germany). This Conference, like the first one, was organized by Faculty of Forestry of the University of Belgrade, IUFRO (specifically by IUFRO Unit: 3.02.00 - Stand Establishment and Treatment), and Reforesta Scientific and professional Society.

Vladan Ivetić

10.07.2018.

Research paper

Variability of beech cupules in Serbia

Beech is the most important tree species in forests of Serbia. However, despite its significance, the taxonomic status of beech in Serbia is unclear. Morphology of the cupules can be successfully used to distinguish species and within species taxa. In this paper we report results on cupules morphology, measuring the same attributes reported by Mišić (1955): 1) Length of the longest valve of cupule, 2) Width of that valve (at widest point), 3) Distance between base of the longest valve and peduncle, 4) Length of cupule without peduncle, and 5) Length of peduncle. The length of cupule and peduncle length from 12 populations in Serbia are in the range reported for Fagus sylvatica in Serbia and western Eurasia. Results of cluster analysis shows a grouping of populations in two groups:1) the southeast group, and 2) group consist of populations from northwest, east and southeast of Serbia. Populations from this southeast group also consist the group of populations on altitude over 850 m, indicating presence of ecotypes. The exception is population from Stara Planina (1,520 m a.s.l.) which is grouped with populations from altitudes under 850 m.

Vladan Ivetić, Ivona Kerkez, Ivan Denić, Jovana Devetaković

10.07.2018.

Research paper

The effect of half-sib lines on morphological attributes of one-year old Fraxinus angustifolia seedlings

Aiming to produce forest reproductive material at the family level, we tested quality of twenty Fraxinus angustifolia Vahl half-sib lines in the nursery experiment. We measured height (HT) and root collar diameter (DIA) of one-year old seedlings, and we monitored germination rate and mortality during the first growing season. Seeds of 20 half-sib lines originating from Sombor area (north Serbia) were sown in seedbed on autumn 2016. The seed dormancy was not broken in all seeds equally, resulting with un-uniform germination over a two months period. All measured attributes shows a statistically significant differences, indicating a strong effect of genetic control. The highest average and DIA were measured in half-sib line 7 (22.62 cm; 5.79 mm, respectively). The lowest average DIA was measured in half-sib lines 18 (2.85 mm) and 19 (2.94 mm), which can be result of growing density effect (104 seedlings m-2 for line 18 and 126 seedlings m-2 for line 19, compared to 40 seedlings m-2 for line 7). The correlation between DIA and HT is strong and positive (R=0.90). Due to its morphological superiority, half-sib line 7 could be recommended for mass production of Fraxinus angustifolia seedlings at the family level.

Ivona Kerkez, Marina Nonić, Jovana Devetaković, Mirjana Šijačić-Nikolić, Vladan Ivetić

10.07.2018.

Review paper

Fertilizer trials for bareroot nurseries in North America

In North America, most tree nursery nutrition publications during the past two decades involved either container-grown stock or stock grown in greenhouses. In contrast, most bareroot nursery fertility trials in North America were published during the last century. As a result, some bareroot fertilization recommendations have remained the same since 1980 and some practices continue to be based on myths and assumptions. The bareroot nursery industry in the USA might benefit if the next generation of graduate students will consider testing old and new theories about nursery fertilization. Hopefully, they will discover new facts so that future fertilization regimes will be based on science. This paper provides various fertilizer trials that should be established in bareroot nurseries.

David B. South

10.07.2018.

Research paper

Breaking seed coat dormancy of six tree species

Breaking physical dormancy in some forest seeds is a challenge for scientists and forest managers to obtain an homogeneous germination for larger seed samples. The role played by the seed coat in seed dormancy of six trees with great interest in agroforestry (Robinia pseudoacacia, Leucaena leucocephala, Erythrostemon gilliesii, Styphnolobium japonicum, Acacia dealbata and Brachychiton populneus) was tested by the effects of the pretreatment and its duration on the performance of seed germination, by considering the final germination percentage (FGP) and the mean germination time (MGT). These parameters are estimated at various times of incubation (5, 10 and 15 days) in Petri dishes and stored in darkness at (25 ± 2°C). The pretreatment consists of an immersion of seeds in concentrated sulphuric acid during 30, 60 and 90 minutes. Sowing without pretreatment (control) revealed no germination induction for B. populneus and A. dealbata, except for R. pseudoacacia, L. leucocephala and S. japonica where the germination does not exceed 12% for all the experience duration. Generally, pretreatments were very useful to improve seed germination. The time of immersion into sulphuric acid significantly affected (P < 0.0001) the FGP and the MGT in all studied species. A duration of 30 minutes of soaking was adequate to give a very high rate of germination for L. leucocephala, E. gilliesii, S. japonica and A. dealbata with respective FGP of 100%, 95%, 100% and 100%. However, an extended duration of pretreatment of 60 minutes was necessary for a maximal germination for R. pseudoacacia and B. populneus with FGP of 85% and 100%, respectively. A prolonged duration of 90 minutes of presowing was very fatal for L. leucocephala, A. dealbata and B. populneus. An excellent germinative strength is characterized by a higher FGP and a reduced MGT.

Abdenour Kheloufi, Lahouaria Mansouri, Nada Aziz, Meriem Sahnoune, Sarra Boukemiche, Boutheina Ababsa